Abstract: Learners that are exposed to the same training data might generalize differently due to differing inductive biases. In neural network models, inductive biases could in theory arise from any aspect of the model architecture. We investigate which architectural factors affect the generalization behavior of neural sequence-to-sequence models trained on two syntactic tasks, English question formation and English tense reinflection. For both tasks, the training set is consistent with a generalization based on hierarchical structure and a generalization based on linear order. All architectural factors that we investigated qualitatively affected how models generalized, including factors with no clear connection to hierarchical structure. For example, LSTMs and GRUs displayed qualitatively different inductive biases. However, the only factor that consistently contributed a hierarchical bias across tasks was the use of a tree-structured model rather than a model with sequential recurrence, suggesting that human-like syntactic generalization requires architectural syntactic structure.